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Bilobed Anther: Structure, Functions, FAQs, Theca

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Bilobed Anther: Structure, Functions, FAQs, Theca

Edited By Team Careers360 | Updated on Jan 29, 2024 04:56 PM IST

The male reproductive organ known as the "Stamen" or "Androecium" of a flower is composed of anthers. It is the primary pollen-producing portion of a sexually active flower and typically consists of two lobes connected by connective tissue. It carries the reproductive components, or male gametes, within the microsporangia (that later form pollen sacs). When these pollen grains germinate, an ovule becomes fertilised.

Male Reproductive Part of Flower

Stamen

The microsporangia are located in the anther, which is positioned on top of the stalk and makes up the stamen. Commonly two-lobed, anthers are joined to the filament at the base or in the centre of the anther. The connective tissue, a protrusion of the filament containing the conducting strands, is the sterile tissue found in the space in between the lobes. It may be seen as an addition on the dorsal side of the anthers. A pollen grain develops from a microspore in the microsporangium and houses the male gametophyte.

The components of the male reproductive system that produce pollen are called anthers. Through the connective tissue, it is joined to the stalk.

The microsporangia are where the pollen is formed. Each microsporangium produces hundreds of microspores. To make room for additional spores, the anthers are lobbed.

Types

The anther is commonly bilobed, but based on the number of lobes, it can be:

  • Unilobed - anther with a single lobe.

  • Tri-lobed - anther with three lobes

There are even anthers with more than three lobes.

Anthers can take one of the following forms depending on their "position of attachment," or the point at which they are joined to the filament of the stamen:

  1. Basifixed: A common kind of attachment in which the filament tip is connected to the anther by "its base."

  2. Dorsifixed: The anther is fixed "dorsally" and almost medially to the filament's tip.

  3. Sub-basifixed: The filament tip is connected to the anther "close to its base."

  4. The versatile anther has all the possibilities open to it, including dorsifixed, basifixed, and sub-basifixed, and is connected "quite loosely" and maintains moving freely at the place of attachment with filament.

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Bilobed Anther

An anther is a bilobed, knob-like structure in terms of morphology, and connective tissue links the two lobes together. An anther has two lobes, four chambers, and a single microsporangium in each of them, as seen through a transverse section. Each lobe has two thecae, meaning an anther is dithecous, and the parenchymatous connective tissue divides the two lobes. The four chambers are divided by the stromium, an external longitudinal groove.

Microspores are found within the thecae or microsporangium. An anther is also known as a "tetra microsporangiate" because it contains four microsporangium, which are enclosed and shielded by an epidermis. The microsporangium matures into "archesporial cells," which are the large, individual cells with dense cytoplasm that maintain the hypodermal state.

Later, the archesporial cells go through a process called "periclinal division," a transverse division that separates the cells into:

  • Peripheral primary parietal cells

  • Inner primary sporogenous cells

Three differentiated layers of cells are formed by periclinal and anticlinal divisions of parietal cells, which also serve to protect the inner sporogenous cells.

  1. Outer endothecium: Parietal cells first go through mitosis and differentiate into outer endothecium, which is active at the time of dehiscence..

  2. Middle layer: Parietal cells continue to divide to create this layer, which typically consists of 1-3 layers of parenchymatous cells.

  3. Tapetum: The third layer (Tapetum) is made up of the outer endothecium and the middle layer. It comprises pyramid-shaped cells that exhibit a radial arrangement around the microsporangium. Tapetum serves as a nutritive layer, which provides nourishment to the growing sporogenous cells.

Pollen Grains

Pollen grains are the main fertilising body of the plant. They carry the male gamete to the female reproductive organ known as ovary, through various mediums like air, water, insects etc.

They are of various shapes and sizes and commonly have a thorny, rough outer covering that is inert in nature. They are produced inside the male sporangia in hundreds of numbers and come out through the process of dehiscence.

Important definitions

  1. Androecium - a group or bunch of stamens of flowers collectively known as Androecium.

  2. Microsporangia - the male gamete producing body inside an anther.

  3. Theca - Theca refers to the anther lobe's cavity. The term "dithecous" refers to lobes with two thecae. There are two pollen sacs, or microsporangia, on each theca.

  4. Archesporial cells : Archesporial cells or archesporia are a type of cell or cell structure found in sporophytes where spores may eventually form during the alternate generation.

Important Events

  • Anthesis: This is the time when the anthers of the developing flower emerge from the spikelet. When a flower reaches maturity, during the stage of anthesis, the process of dehiscence takes place.

  • Dehiscence: When an anther reaches maturity, it explodes and releases pollen grains. As a result, the process of an anther dehiscence is what causes pollen to be released. Pollen grains are released by an anther depending on the direction of dehiscence.

Frequently Asked Question (FAQs)

1. Is anther haploid or diploid?

An anther is diploid (2n) in structure, along with the plant itself.

2. What is the process of cutting down an anther?

Emasculation is the removal of stamens from the female parents' bisexual flowers in order to prevent self-pollination. It is done before the anthers are mature.

3. Does one female flower possess another?

No, an anther is a male gamete-producing body, so unless the flower is not bisexual or male, it lacks an anther.

4. Anthers are important for self-pollination.

Yes, both the anther and the ovary must be present in the same flower, or at least in different flowers of the same plant, to achieve self-pollination.

5. What if we remove the anthers before their dehiscence?

Removing the anthers before they dehisce will result in the death of both the anthers and the pollen within them. It is a practice that is used to prevent self-pollination.

Bilobed Anther: Structure, Functions, FAQs, Theca

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